But birds don't have heel bones. they don't have an calcaneus.
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In the picture above, the raven prepares to make contact heel first with toes pointed upwards. (the raven in the SS is running with a recently updated animation, IMO it looks good enough. But during walking, the problem is much more pronounced.)
But in birds, the toes contact the ground first. Every motion of the swinging foot, the direction of the toes point to set up so that contact is made by the toes first (toes point towards the ground, during swing. Unlike humans, where toes point toward sky during swing)
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In both of the above figures, there is no moment in which the wingless bird(or any bird)'s toes point towrads the sky, because it is always pointing down to prepare to touch ground first.
Millions of years of evolution and probably some physics have said that this is the most practical and efficient way to use those joints.
Edited by Tennex, 13 January 2013 - 09:13 AM.